Suzume, 17, lost her mother as a little girl. On her way to school, she meets a mysterious young man. But her curiosity unleashes a calamity that endangers the entire population of Japan, and so Suzume embarks on a journey to set things right.
The season is fall and the city of Neo-Venezia is covered in fallen leaves. Something is bothering Anya, who works at Orange Planet. Her senior colleagues Alice and Athena haven't seen each other due to their busy schedules, which is making Athena feel sad. But for some reason, Alice seems to be avoiding her altogether. With her friends Ai and Azusa's help, Anya tries to find a definitive way for Alice and Athena to get together. But during this time, Anya learns of a situation that only she can see because of where she stands.
In a world bursting at the seams with moe monsters and humanoids of the horned sort, which brave heroes will take it upon themselves to review the beastly babes of the red-light district? Can only one be crowned the ultimate title of best girl? Behold the most tantalizing of trials.